Tethysaurus

Tethysaurus
Temporal range: Late Cretaceous, 93–89 Ma
Tethysaurus skeleton
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Order: Squamata
Family: Mosasauridae
Subfamily: Tethysaurinae
Genus: Tethysaurus
Bardet, Pereda-Suberbiola & Jalil, 2003
Type species
Tethysaurus nopscai
Bardet, Pereda-Suberbiola & Jalil, 2003

Tethysaurus is an extinct genus of tethysaurine mosasauroid from the Early Turonian (Late Cretaceous) period. The only species is Tethysaurus nopscai.

Discovery

Fossil

The name means "Tethys' lizard of Nopsca", a reference to the Greek goddess of the sea Tethys (also the name of the Tethys Ocean, an ancient sea between southern Europe and northern Africa) and to the Hungarian paleontologist Baron Ferenc Nopsca, who made pioneering studies on Adriatic aquatic squamates. It was found near the villages of Tadirhourst and Asfla in the region of Goulmima, Errachidia Province, in Morocco, with three referred specimens that included a nearly complete articulated skull, mandible, vertebrae and portions of the appendicular skeleton. The diagnosis after Bardet et al. is "(...) prefrontal strongly vaulted in anterior view; parietal exhibits a triangular table ending posteriorly in two pointed pegs overlying the supraoccipital; jugal with a large and wide ascending ramus; medullar floor of the basioccipital pierced by three foramina; splenial with a large notched dorsomedial process; surangular exposed medially ventral to the coronoid; dental formula: 19-20 maxillary, 15-19 pterygoid and at least 19 dentary teeth; large paracotylar and parazygosphenal foramina on vertebrae."[1]

Description

An average-sized mosasaur of 3 m (9.8 ft) length, Tethysaurus displays a number of basal and derived features that led to an initial classification as an intermediate stage between primitive aigialosaurids of the Cenomanian and derived mosasaurids from the Turonian to the Maastrichtian.[2] More recent analysis put Tethysaurus in a clade along Russellosaurus and Yaguarasaurus called the parafamily Russellosaurina, as a basal Turonian clade of Mosasauridae.[3]

Phylogeny

Tethysaurus sclerotic eye ring

Cladogram based in the analysis by Makádi et al. in 2012:[4]

Tethysaurinae

Pannoniasaurus inexpectatus




Tethysaurus nopcsai




Yaguarasaurus columbianus



Russellosaurus coheni





See also

References

  1. Nathalie Bardet, Xabier Pereda Suberbiola, and Nour-Eddine Jalil. A new mosasauroid (Squamata) from the Late Cretaceous (Turonian) of Morocco. Comptes Rendus Palevol, Volume 2, Issue 8, December 2003, page 611.
  2. Nathalie Bardet, Xabier Pereda Suberbiola, and Nour-Eddine Jalil. A new mosasauroid (Squamata) from the Late Cretaceous (Turonian) of Morocco. Comptes Rendus Palevol, Volume 2, Issue 8, December 2003, Pages 607-616.
  3. Polcyn, M. J. and Bell, G. L., Jr. 2005. Russellosaurus coheni n. gen., n. sp., a 92 million-year-old mosasaur from Texas (USA), and the definition of the parafamily Russellosaurina. Netherlands Journal of Geosciences 84(3):321-333.
  4. Makádi, L. S.; Caldwell, M. W.; Ősi, A. (2012). Butler, Richard J, ed. "The First Freshwater Mosasauroid (Upper Cretaceous, Hungary) and a New Clade of Basal Mosasauroids". PLoS ONE. 7 (12): e51781. doi:10.1371/journal.pone.0051781.


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