List of ant genera

The following is a list of worldwide ant genera.

Contents:
Agroecomyrmecinae • Amblyoponinae • Aneuretinae • Armaniinae • Brownimeciinae • Dolichoderinae • Dorylinae • Ectatomminae • Formiciinae • Formicinae • Heteroponerinae • Leptanillinae • Martialinae • Myrmeciinae • Myrmicinae • Paraponerinae • Ponerinae • Proceratiinae • Pseudomyrmecinae • Sphecomyrminae • Incertae sedis • Excluded from Formicidae •
See also        Notes        References

Key

Genus name	The scientific name of the genus. Extinct taxa are indicated by a †.
Binomial authority	The binomial authority—the person who first described the species using an available scientific name.
Classified	The year the genus was formally described and classified.
No. of species	The number of species, both extant and extinct, that are members of the genus.
Type species	The type species that was designated by the original author, unless noted otherwise. Extinct taxa are indicated by a †.
Example image	An image (if available) of a species who is a member of the genus.
References	Most citations provided are the first publications that describe the genus; additional citations are added if the original source does not contain all provided information.

Subfamilies

Agroecomyrmecinae

The subfamily Agroecomyrmecinae represents two extant genera and two fossil genera, once widespread in both hemispheres during the early Tertiary.^[1][2] The subfamily was originally classified as Agroecomyrmecini, a Myrmicinae tribe until English myrmecologist Barry Bolton raised the tribe to subfamily status in 2003.^[3][4] Most specimens collected are from Central America and Mexico, although one type specimen of an Agroecomyrmecinae species was collected from Ghana.^[5]

Subfamily Agroecomyrmecinae Carpenter, 1930 – four genera, five species[1]
Genus name	Binomial authority	Classified	No. of species	Type species	Example image	Ref(s)
Agroecomyrmex !†Agroecomyrmex	Wheeler	1910	1	Myrmica duisburgi !†Myrmica duisburgi	†Agroecomyrmex duisburgi	[6][7]
Ankylomyrma	Bolton	1973	1	Ankylomyrma coronacantha	Ankylomyrma coronacantha	[8]
Eulithomyrmex !†Eulithomyrmex	Carpenter	1935	2	Lithomyrmex rugosus !†Lithomyrmex rugosus	†Eulithomyrmex rugosus	[9]
Tatuidris	Brown & Kempf	1968	1	Tatuidris tatusia	Tatuidris tatusia	[2]

Amblyoponinae

The subfamily Amblyoponinae represents 10 extant genera and one fossil genus. Established by Swiss myrmecologist Auguste Forel in 1893, these ants are specialist predators, distributed worldwide in the tropics.^[10][11][12]

Subfamily Amblyoponinae Forel, 1893 – 11 genera, 146 species[13]
Genus name	Binomial authority	Classified	No. of species	Type species	Example image	Ref(s)
Adetomyrma	Ward	1994	9	Adetomyrma venatrix	Adetomyrma venatrix	[14]
Amblyopone	Erichson	1842	10	Amblyopone australis	Amblyopone australis	[15]
Apomyrma	Brown, Gotwald & Levieux	1970	1	Apomyrma stygia	Apomyrma stygia	[16]
Casaleia !†Casaleia	Pagliano & Scaramozzino	1990	4	Protamblyopone inversa !†Protamblyopone inversa	†Casaleia eocenica	[17]
Fulakora	Mann	1919	25	Stigmatomma celata	Fulakora celata	[18][19]
Myopopone	Roger	1861	2	Myopopone maculata	Myopopone castanea	[20][21]
Mystrium	Roger	1862	14	Mystrium mysticum	Mystrium mysticum	[22]
Onychomyrmex	Emery	1895	4	Onychomyrmex hedleyi	Onychomyrmex hedleyi	[23]
Prionopelta	Mayr	1866	22	Prionopelta punctulata	Prionopelta punctulata	[24]
Stigmatomma	Roger	1859	54	Stigmatomma denticulatum	Stigmatomma denticulatum	[25]
Xymmer	Santschi	1914	1	Stigmatomma muticum	Xymmer muticus	[26]

Aneuretinae

The subfamily Aneuretinae represents one extant genus and eight fossil genera established by Italian entomologist Carlo Emery in 1913. Only a single species of this subfamily is extant, the Sri Lankan relict ant (Aneuretus simoni), endemic to Sri Lanka.^[27]

Subfamily Aneuretinae Emery, 1913 – nine genera, 12 species[28]
Genus name	Binomial authority	Classified	No. of species	Type species	Example image	Ref(s)
Aneuretellus !†Aneuretellus	Dlussky	1988	1	Aneuretellus deformis !†Aneuretellus deformis	N/A	[29]
Aneuretus	Emery	1893	1	Aneuretus simoni	Aneuretus simoni	[30]
Britaneuretus !†Britaneuretus	Dlussky & Perfilieva	2014	1	Britaneuretus anglicus !†Britaneuretus anglicus	N/A	[31]
Burmomyrma !†Burmomyrma	Dlussky	1996	1	Burmomyrma rossi !†Burmomyrma rossi	†Burmomyrma rossi	[32]
Cananeuretus !†Cananeuretus	Engel & Grimaldi	2005	1	Cananeuretus occidentalis !†Cananeuretus occidentalis	N/A	[33]
Mianeuretus !†Mianeuretus	Carpenter	1930	2	Mianeuretus mirabilis !†Mianeuretus mirabilis	†Mianeuretus mirabilis	[3]
Paraneuretus !†Paraneuretus	Wheeler	1915	3	Paraneuretus tornquisti !†Paraneuretus tornquisti	†Paraneuretus tornquisti	[7]
Pityomyrmex !†Pityomyrmex	Wheeler	1915	1	Pityomyrmex tornquisti !†Pityomyrmex tornquisti	†Pityomyrmex tornquisti	[7]
Protaneuretus !†Protaneuretus	Wheeler	1915	1	Protaneuretus succineus !†Protaneuretus succineus	†Protaneuretus succineus	[7]

Armaniinae

The subfamily Armaniinae (or family Armaniidae) contains seven extinct genera from a series of Cretacous fossils found in Asia and Africa.^[34] These ants are treated as a group of "ant-like wasps",^[33] and they appear to lack a metapleural gland which would exclude them from the family Formicidae. However, the lack of a metapleural gland could be due to poorer preservation.^[35]

Subfamily Armaniinae Dlussky, 1983 – seven genera, 14 species
Genus name	Binomial authority	Classified	No. of species	Type species	Example image	Ref(s)
Archaeopone !†Archaeopone	Dlussky	1975	2	Archaeopone kzylzharica !†Archaeopone kzylzharica	N/A	[36]
Armania !†Armania	Dlussky	1983	4	Armania robusta !†Armania robusta	N/A	[37]
Dolichomyrma !†Dolichomyrma	Dlussky	1975	2	Dolichomyrma longiceps !†Dolichomyrma longiceps	N/A	[36]
Khetania !†Khetania	Dlussky	1999	1	Khetania mandibulata !†Khetania mandibulata	N/A	[38]
Orapia !†Orapia	Dlussky, Brothers & Rasnitsyn	2004	2	Orapia rayneri !†Orapia rayneri	†Orapia rayneri	[39]
Poneropterus !†Poneropterus	Dlussky	1983	1	Poneropterus sphecoides !†Poneropterus sphecoides	N/A	[37]
Pseudarmania !†Pseudarmania	Dlussky	1983	2	Pseudarmania rasnitsyni !†Pseudarmania rasnitsyni	N/A	[37]

Brownimeciinae

The subfamily Brownimeciinae contains the single genus Brownimecia which only has one species, Brownimecia clavata. It was described in 1997 after a fossilised specimen was collected from Cretaceous amber from New Jersey, and was initially placed in the subfamily Ponerinae.^[34] The species was later classified into its own subfamily in 2003 by Barry Bolton.^[4]

Subfamily †Brownimeciinae Bolton, 2003 – one genus, one species[40]
Genus name	Binomial authority	Classified	No. of species	Type species	Example image	Ref(s)
Brownimecia !†Brownimecia	Grimaldi, Agosti & Carpenter	1997	1	Brownimecia clavata !†Brownimecia clavata	†Brownimecia clavata	[34]

Dolichoderinae

The subfamily Dolichoderinae was established by Forel in 1878, which represents 28 extant genera and 20 fossil genera. The subfamily presents a great diversity of species throughout the world, mainly in the tropics. Most species are generalised scavengers, but some are predacious.^[41][42]

Subfamily Dolichoderinae Forel, 1878 – 48 genera, 843 species[43]
Genus name	Binomial authority	Classified	No. of species	Type species	Example image	Ref(s)
Alloiomma !†Alloiomma	Zhang	1989	2	Alloiomma changweiensis !†Alloiomma changweiensis	N/A	[44]
Anillidris	Santschi	1936	1	Anillidris bruchi	Anillidris bruchi	[45]
Anonychomyrma	Donisthorpe	1947	26	Anonychomyrma myrmex	Anonychomyrma gilberti	[46]
Aptinoma	Fisher	2009	2	Aptinoma mangabe	Aptinoma mangabe	[47]
Arnoldius	Dubovikoff	2005	3	Bothriomyrmex flavus	Arnoldius flavus	[48]
Asymphylomyrmex !†Asymphylomyrmex	Wheeler	1915	1	Asymphylomyrmex balticus !†Asymphylomyrmex balticus	†Asymphylomyrmex balticus	[7]
Axinidris	Weber	1941	21	Axinidris acholli	Axinidris acholli	[49]
Azteca	Forel	1878	86	Liometopum xanthochroum	Azteca xanthochroum	[50]
Bothriomyrmex	Emery	1869	24	Bothriomyrmex costae	Bothriomyrmex costae	[51]
Chronomyrmex !†Chronomyrmex	McKellar, Glasier & Engel	2013	1	Chronomyrmex medicinehatensis !†Chronomyrmex medicinehatensis	†Chronomyrmex medicinehatensis	[52]
Chronoxenus	Santschi	1919	6	Bothriomyrmex myops	Chronoxenus myops	[53]
Ctenobethylus !†Ctenobethylus	Brues	1939	1	Ctenobethylus succinalis !†Ctenobethylus succinalis	†Ctenobethylus goepperti	[54]
Doleromyrma	Forel	1907	2	Tapinoma darwinianum	Doleromyrma darwiniana	[55]
Dolichoderus	Lund	1831	181	Formica attelaboides	Dolichoderus attelaboides	[56]
Dorymyrmex	Mayr	1866	60	Dorymyrmex flavescens	Dorymyrmex flavescens	[24]
Ecphorella	Forel	1909	1	Tapinoma wellmani	Ecphorella wellmani	[57]
Elaeomyrmex !†Elaeomyrmex	Carpenter	1930	2	Elaeomyrmex gracilis !†Elaeomyrmex gracilis	†Elaeomyrmex gracilis	[3]
Elaphrodites !†Elaphrodites	Zhang	1989	2	Elaphrodites scutulatus !†Elaphrodites scutulatus	N/A	[44]
Eldermyrmex !†Eldermyrmex	Heterick & Shattuck	2011	1	Eldermyrmex oblongiceps !†Iridomyrmex oblongiceps	†Eldermyrmex oblongiceps	[58]
Emplastus !†Emplastus	Donisthorpe	1920	12	Emplastus emeryi !†Emplastus emeryi	†Emplastus britannicus	[59]
Eotapinoma !†Eotapinoma	Dlussky	1988	3	Eotapinoma gracilis !†Eotapinoma gracilis	N/A	[29]
Eurymyrmex !†Eurymyrmex	Zhang, Sun & Zhang	1994	1	Eurymyrmex geologicus !†Eurymyrmex geologicus	N/A	[60]
Forelius	Emery	1888	18	Iridomyrmex mccooki	Forelius mccooki	[61]
Froggattella	Forel	1902	2	Acantholepis kirbii !Acantholepis kirbii	Froggattella kirbii	[62]
Gracilidris	Wild & Cuezzo	2006	2	Gracilidris pombero	Gracilidris pombero	[63]
Iridomyrmex	Mayr	1862	84	Formica detecta	Iridomyrmex purpureus	[21][64]
Kotshkorkia !†Kotshkorkia	Dlussky	1981	1	Kotshkorkia laticeps !†Kotshkorkia laticeps	N/A	[65]
Ktunaxia !†Ktunaxia	Lapolla & Greenwalt	2015	1	Ktunaxia jucunda !†Ktunaxia jucunda	†Ktunaxia jucunda	[66]
Leptomyrmex	Mayr	1862	29	Formica erythrocephala	Leptomyrmex erythrocephalus	[64]
Leptomyrmula !†Leptomyrmula	Emery	1913	1	Leptomyrmula maravignae !†Leptomyrmula maravignae	†Leptomyrmula maravignae	[67]
Linepithema	Mayr	1866	20	Linepithema fuscum	Linepithema fuscum	[24]
Liometopum	Mayr	1861	28	Liometopum microcephalum	Liometopum microcephalum	[68]
Loweriella	Shattuck	1992	1	Loweriella boltoni	Loweriella boltoni	[69]
Miomyrmex !†Miomyrmex	Carpenter	1930	2	Miomyrmex impactus !†Miomyrmex impactus	†Miomyrmex impactus	[3]
Nebothriomyrmex	Dubovikov	2004	1	Nebothriomyrmex majeri	Nebothriomyrmex majeri	[70]
Ochetellus	Shattuck	1992	7	Ochetellus glaber	Ochetellus glaber	[71]
Papyrius	Shattuck	1992	2	Papyrius nitidus	Papyrius nitidus	[71]
Petraeomyrmex !†Petraeomyrmex	Carpenter	1930	1	Petraeomyrmex minimus !†Petraeomyrmex minimus	†Petraeomyrmex minimus	[3]
Philidris	Shattuck	1992	9	Philidris cordata	Philidris cordata	[71]
Proiridomyrmex !†Proiridomyrmex	Dlussky & Rasnitsyn	2003	2	Proiridomyrmex vetulus !†Proiridomyrmex vetulus	†Proiridomyrmex rotundatus	[66]
Protazteca !†Protazteca	Carpenter	1930	5	Protazteca elongata !†Protazteca elongata	†Protazteca elongata	[3]
Ravavy	Fisher	2009	1	Ravavy miafina	Ravavy miafina	[47]
Tapinoma	Förster	1850	74	Tapinoma erraticum	Tapinoma erraticum	[72]
Technomyrmex	Mayr	1872	97	Technomyrmex strenuus	Technomyrmex strenuus	[73]
Turneria	Forel	1895	8	Turneria bidentata	Turneria bidentata	[74]
Usomyrma !†Usomyrma	Dlussky et al.	2014	1	Usomyrma mirabilis !†Usomyrma mirabilis	†Usomyrma mirabilis	[75]
Yantaromyrmex !†Yantaromyrmex	Dlussky & Dubovikoff	2013	5	Yantaromyrmex geinitzi !†Yantaromyrmex geinitzi	†Yantaromyrmex geinitzi	[76]
Zherichinius !†Zherichinius	Dlussky	1988	2	Zherichinius horribilis !†Zherichinius horribilis	†Zherichinius horribilis	[29]

Dorylinae

The subfamily Dorylinae was established by Leach in 1815, which represents 27 extant genera and one fossil genus. Many species of ant in this subfamily are known as army ants that are distributed in the Old World and New World.^[77][78]

Subfamily Dorylinae Leach, 1815 – 28 genera, 693 species[78][79]
Genus name	Binomial authority	Classified	No. of species	Type species	Example image	Ref(s)
Acanthostichus	Mayr	1887	24	Acanthostichus serratulus	Acanthostichus serratulus	[80]
Aenictogiton	Emery	1901	7	Aenictogiton fossiceps	Aenictogiton fossiceps	[81]
Aenictus	Shuckard	1840	184	Aenictus ambiguus	Aenictus ambiguus	[82]
Cerapachys	Smith	1857	5	Cerapachys antennatus	Cerapachys antennatus	[21][83]
Cheliomyrmex	Mayr	1870	4	Cheliomyrmex morosus	Cheliomyrmex morosus	[84]
Chrysapace	Crawley	1924	3[lower-alpha 1]	Chrysapace jacobsonii	Chrysapace jacobsonii	[85]
Cylindromyrmex	Mayr	1870	13	Cylindromyrmex striatus	Cylindromyrmex striatus	[84]
Dorylus	Fabricius	1793	60	Dorylus helvolus	Dorylus helvolus	[86]
Eburopone	Borowiec	2016	1	Cerapachys wroughtoni	Eburopone wroughtoni	[78]
Eciton	Latreille	1804	12	Eciton hamatum	Eciton hamatum	[87][88]
Eusphinctus	Emery	1893	2	Eusphinctus furcatus	Eusphinctus furcatus	[30]
Labidus	Jurine	1807	7	Labidus coecus	Labidus coecus	[89]
Leptanilloides	Mann	1923	19	Leptanilloides biconstricta	Leptanilloides biconstricta	[90]
Lioponera	Mayr	1879	73	Lioponera longitarsus	Lioponera longitarsus	[91]
Lividopone	Bolton & Fisher	2016	1	Cerapachys lividus	Lividopone livida	[92]
Neivamyrmex	Borgmeier	1940	128	Neivamyrmex nigrescens	Neivamyrmex nigrescens	[93]
Neocerapachys	Borowiec	2016	2	Cerapachys neotropicus	Neocerapachys neotropicus	[78]
Nomamyrmex	Borgmeier	1936	2	Nomamyrmex esenbeckii	Nomamyrmex esenbeckii	[94]
Ooceraea	Roger	1862	11	Ooceraea fragosa	Ooceraea fragosa	[22]
Parasyscia	Emery, in André	1882	50	Parasyscia piochardi	Parasyscia piochardi	[95]
Procerapachys !†Procerapachys	Wheeler	1915	3	Procerapachys annosus !†Procerapachys annosus	†Procerapachys annosus	[7]
Simopone	Forel	1891	39	Simopone grandidieri	Simopone grandidieri	[96]
Sphinctomyrmex	Mayr	1866	3	Sphinctomyrmex stali	Sphinctomyrmex stali	[97]
Syscia	Roger	1861	5	Syscia typhla	Syscia typhla	[20]
Tanipone	Bolton & Fisher	2012	10	Tanipone hirsuta	Tanipone hirsuta	[98]
Vicinopone	Bolton & Fisher	2012	1	Vicinopone conciliatrix	Vicinopone conciliatrix	[98]
Yunodorylus	Xu	2000	4	Yunodorylus sexspinus	Yunodorylus sexspinus	[99]
Zasphinctus	Wheeler	1918	20	Sphinctomyrmex turneri	Zasphinctus turneri	[100]

Ectatomminae

The subfamily Ectatomminae represents four extant genera and three fossil genera, established in 1895 by Carlo Emery. They are distributed in tropical and warm climates in the New World and Old World, as well as the Indo-Australian regions.^[42]

Subfamily Ectatomminae Emery, 1895 – seven genera, 278 species[101]
Genus name	Binomial authority	Classified	No. of species	Type species	Example image	Ref(s)
Canapone !†Canapone	Dlussky	1999	1	Canapone dentata !†Canapone dentata	N/A	[102]
Ectatomma	Smith	1858	16	Ectatomma tuberculatum	Ectatomma tuberculatum	[21][103]
Electroponera !†Electroponera	Wheeler	1915	1	Electroponera dubia !†Electroponera dubia	†Electroponera dubia	[7]
Gnamptogenys	Roger	1863	143	Gnamptogenys sulcata	Gnamptogenys sulcata	[104][105]
Pseudectatomma !†Pseudectatomma	Dlussky & Wedman	2012	2	Pseudectatomma eocenica !†Pseudectatomma eocenica	†Pseudectatomma eocenica	[11]
Rhytidoponera	Mayr	1862	108	Rhytidoponera araneoides	Rhytidoponera araneoides	[64][105]
Typhlomyrmex	Mayr	1862	7	Scyphodon anomalum	Scyphodon anomalum	[64]

Formiciinae

The subfamily Formiciinae represents one extinct genus of ants dating back to the Eocene.^{[lower-alpha 2]} Ants of the genus Titanomyrma are the largest ants ever known, with queen specimens the size of small hummingbirds. Fossils have been collected from the state of Wyoming and in Germany.^[107][108]

Subfamily †Formiciinae Lutz, 1986 – one, six species[109]
Genus name	Binomial authority	Classified	No. of species	Type species	Example image	Ref(s)
Formicium !†Formicium[lower-alpha 2]	Westwood	1854	3	None[lower-alpha 3]	N/A	[110]
Titanomyrma !†Titanomyrma	Archibald, Johnson, Mathewes & Greenwood	2011	3	Titanomyrma gigantea !†Titanomyrma gigantea	†Titanomyrma gigantea	[107]

Formicinae

The subfamily Formicinae represents 51 extant genera and 30 fossil genera that are globally distributed. Established by French zoologist Pierre André Latreille in 1809, the subfamily has more than 3,000 described species, placing it as the second largest ant subfamily. Despite this, the hyperdiverse genus Camponotus is the most diverse group of ants in the world, with more than 1,100 species described.^[42][111]

Subfamily Formicinae Latreille, 1809 – 81 genera, 3,228 species[112]
Genus name	Binomial authority	Classified	No. of species	Type species	Example image	Ref(s)
Acropyga	Roger	1862	41	Acropyga acutiventris	Acropyga acutiventris	[22]
Agraulomyrmex	Prins	1983	2	Agraulomyrmex meridionalis	Agraulomyrmex meridionalis	[113]
Alloformica	Dlussky	1969	4	Alloformica aberrans	Alloformica aberrans	[114]
Anoplolepis	Santschi	1914	9	Anoplolepis gracilipes	Anoplolepis gracilipes	[115]
Aphomomyrmex	Emery	1899	1	Aphomomyrmex afer	Aphomomyrmex afer	[116]
Bajcaridris	Agosti	1994	3	Bajcaridris theryi	Bajcaridris theryi	[117]
Brachymyrmex	Mayr	1868	44	Brachymyrmex patagonicus	Brachymyrmex patagonicus	[118]
Bregmatomyrma	Wheeler	1929	1	Bregmatomyrma carnosa	Bregmatomyrma carnosa	[119]
Calomyrmex	Emery	1895	9	Calomyrmex laevissimus	Calomyrmex laevissimus	[120]
Camponotites !†Camponotites	Steinbach	1967	5	Camponotites silvestris !†Camponotites silvestris	†Camponotites silvestris	[121]
Camponotus	Mayr	1861	1131	Camponotus ligniperda	Camponotus ligniperda	[21][68]
Cataglyphis	Förster	1850	91	Cataglyphis bicolor	Cataglyphis bicolor	[122]
Cataglyphoides !†Cataglyphoides	Dlussky	2008	2	Cataglyphoides constrictus !†Cataglyphoides constrictus	†Cataglyphoides constrictus	[123]
Chimaeromyrma !†Chimaeromyrma	Dlussky	1988	1	Chimaeromyrma brachycephala !†Chimaeromyrma brachycephala	N/A	[29]
Cladomyrma	Wheeler	1920	13	Cladomyrma hewitti	Cladomyrma hewitti	[124]
Conoformica !†Conoformica	Dlussky	2008	1	Conoformica bitterfeldiana !†Conoformica bitterfeldiana	N/A	[123]
Curtipalpulus !†Curtipalpulus	Hong	2002	1	Curtipalpulus eocenicus !†Curtipalpulus eocenicus	N/A	[125]
Drymomyrmex !†Drymomyrmex	Wheeler	1915	2	Drymomyrmex fuscipennis !†Drymomyrmex fuscipennis	†Drymomyrmex fuscipennis	[7]
Echinopla	Smith	1857	33	Echinopla melanarctos	Echinopla melanarctos	[83][126]
Eoleptocerites !†Eoleptocerites	Hong	2002	2	Eoleptocerites chinensis !†Eoleptocerites chinensis	N/A	[125]
Euprenolepis	Emery	1906	8	Euprenolepis procera	Euprenolepis procera	[127]
Eurytarsites !†Eurytarsites	Hong	2002	1	Eurytarsites fushunensisa !†Eurytarsites fushunensisa	N/A	[125]
Forelophilus	Kutter	1931	3	Forelophilus overbecki	Forelophilus overbecki	[128]
Formica	Linnaeus	1758	234	Formica rufa	Formica rufa	[129]
Fushuniformica !†Fushuniformica	Hong	2002	1	Fushuniformica spinata !†Fushuniformica spinata	N/A	[125]
Gesomyrmex	Mayr	1868	17	Gesomyrmex hoernesi !†Gesomyrmex hoernesi	†Gesomyrmex hoernesi	[130]
Gigantiops	Roger	1863	1	Gigantiops destructor	Gigantiops destructor	[131]
Glaphyromyrmex !†Glaphyromyrmex	Wheeler	1915	1	Glaphyromyrmex oligocenicus !†Glaphyromyrmex oligocenicus	†Glaphyromyrmex oligocenicus	[7]
Heeridris !†Heeridris	Dlussky & Putyatina	2014	1	Heeridris croaticus !†Heeridris croaticus	N/A	[132]
Huaxiaformica !†Huaxiaformica	Hong	2002	1	Huaxiaformica mallepetiolata !†Huaxiaformica mallepetiolata	N/A	[125]
Iberoformica	Tinaut	1990	2	Iberoformica subrufa	Iberoformica subrufa	[133]
Imhoffia !†Imhoffia	Heer	1850	2	Imhoffia nigra !†Imhoffia nigra	N/A	[134]
Kyromyrma !†Kyromyrma	Grimaldi & Agosti	2000	1	Kyromyrma neffi !†Kyromyrma neffi	†Kyromyrma neffi	[135]
Lasiophanes	Emery	1895	6	Lasiophanes atriventris	Lasiophanes atriventris	[136]
Lasius	Fabricius	1804	134	Formica nigra !†Formica nigra	†Formica nigra	[137]
Lepisiota	Santschi	1926	81	Lepisiota rothneyi	Lepisiota rothneyi	[138]
Leptogasteritus !†Leptogasteritus	Hong	2002	1	Leptogasteritus capricornutes !†Leptogasteritus capricornutes	N/A	[125]
Leucotaphus !†Leucotaphus	Donisthorpe	1920	2	Leucotaphus gurnetensis !†Leucotaphus gurnetensiss	†Leucotaphus gurnetensiss	[59]
Liaoformica !†Liaoformica	Hong	2002	1	Liaoformica longipetiolata !†Liaoformica longipetiolata	N/A	[125]
Longiformica !†Longiformica	Hong	2002	2	Longiformica dongzhouheensis !†Longiformica dongzhouheensis	N/A	[125]
Magnogasterites !†Magnogasterites	Hong	2002	1	Magnogasterites curticornutus !†Magnogasterites curticornutus	N/A	[125]
Melophorus	Lubbock	1883	23	Melophorus bagoti	Melophorus bagoti	[139]
Myrmecocystus	Wesmael	1838	29	Myrmecocystus mexicanus	Myrmecocystus mexicanus	[140]
Myrmecorhynchus	André	1896	3	Myrmecorhynchus emeryi	Myrmecorhynchus emeryi	[141]
Myrmelachista	Roger	1863	56	Myrmelachista kraatzii	Myrmelachista kraatzii	[104]
Myrmoteras	Forel	1893	41	Myrmoteras binghamii	Myrmoteras binghamii	[142]
Notoncus	Emery	1895	6	Notoncus ectatommoide	Notoncus ectatommoide	[23]
Notostigma	Emery	1920	2	Notostigma carazzii	Notostigma carazzii	[143]
Nylanderia	Emery	1906	110	Nylanderia vividula	Nylanderia vividula	[127]
Oecophylla	Smith	1860	15	Oecophylla smaragdina	Oecophylla smaragdina	[144]
Opisthopsis	Dalla Torre	1893	13	Myrmecopsis respiciens	Opisthopsis respiciens	[145]
Orbicapitia !†Orbicapitia	Hong	2002	1	Orbicapitia reticulata !†Orbicapitia reticulata	N/A	[125]
Ovalicapito !†Ovalicapito	Hong	2002	1	Ovalicapito fushunensis !†Ovalicapito fushunensis	N/A	[125]
Ovaligastrula !†Ovaligastrula	Hong	2002	1	Ovaligastrula xilutianensis !†Ovaligastrula xilutianensis	N/A	[125]
Overbeckia	Viehmeyer	1916	1	Overbeckia subclavata	Overbeckia subclavata	[146]
Paraparatrechina	Donisthorpe	1947	36	Paratrechina pallida	Paraparatrechina pallida	[147]
Paratrechina	Motschoulsky	1863	5	Paratrechina currens	Paratrechina longicornis	[126][148]
Petalomyrmex	Snelling	1979	1	Petalomyrmex phylax	Petalomyrmex phylax	[149]
Phasmomyrmex	Stitz	1910	4	Phasmomyrmex sericeus	Phasmomyrmex buchneri	[150]
Plagiolepis	Mayr	1861	72	Formica pygmaea	Plagiolepis pygmaea	[68]
Polyergus	Latreille	1804	14	Formica rufescens	Polyergus rufescens	[87]
Polyrhachis	Smith	1857	698	Formica bihamata	Polyrhachis bihamata	[83]
Prenolepis	Mayr	1861	17	Tapinoma nitens	Prenolepis nitens	[21][68]
Prodimorphomyrmex !†Prodimorphomyrmex	Wheeler	1915	1	Prodimorphomyrmex !†Prodimorphomyrmex primigenius	†Prodimorphomyrmex primigenius	[7]
Proformica	Ruzsky	1902	25	Formica nasuta	Proformica nasuta	[151]
Prolasius	Forel	1892	19	Formica advena	Prolasius advena	[152]
†Protoformica	Dlussky	1967	1	Protoformica !†Formica proformicoides	N/A	[153]
Protrechina !†Protrechina	Wilson	1985	1	Protrechina carpenteri !†Protrechina carpenteri	†Protrechina carpenteri	[154]
Pseudocamponotus !†Pseudocamponotus	Carpenter	1930	1	Pseudocamponotus elkoanus !†Pseudocamponotus elkoanus	†Pseudocamponotus elkoanus	[3]
Pseudolasius	Emery	1887	50	Formica familiaris	Pseudolasius familiaris	[21][155]
Pseudonotoncus	Clark	1934	2	Pseudonotoncus hirsutus	Pseudonotoncus hirsutus	[156]
Rossomyrmex	Arnol'di	1928	4	Rossomyrmex proformicarum	Rossomyrmex proformicarum	[157]
Santschiella	Forel	1916	1	Santschiella kohli	Santschiella kohli	[158]
Sicilomyrmex !†Sicilomyrmex	Wheeler	1915	1	Pseudocamponotus elkoanus !†Gesomyrmex corniger	†Sicilomyrmex corniger	[7]
Sinoformica !†Sinoformica	Hong	2002	1	Sinoformica longicapitata !†Sinoformica longicapitata	N/A	[125]
Sinotenuicapito !†Sinotenuicapito	Hong	2002	1	Sinotenuicapito badis !†Sinotenuicapito badis	N/A	[125]
Stigmacros	Forel	1905	49	Acantholepis froggatti	Stigmacros froggatti	[159]
Tapinolepis	Emery	1925	14	Plagiolepis tumidula	Tapinolepis tumidula	[160]
Teratomyrmex	McAreavey	1957	3	Teratomyrmex greavesi	Teratomyrmex greavesi	[161]
Wilsonia !†Wilsonia	Hong	2002	2	Wilsonia megagastrosa !†Wilsonia megagastrosa	N/A	[125]
Zatania	LaPolla, Kallal & Brady	2012	6	Paratrechina cisipa	Zatania albimaculata	[162]

Heteroponerinae

The subfamily Heteroponerinae represents three extant genera of ants, established in 2003 when Barry Bolton divided the Ponerinae subfamily into six subfamilies.^[42] These ants are known from the Neotropics of Central America and South America while Aulacopone relicta is from Azerbaijan.^[163]

Subfamily Heteroponerinae Bolton, 2003 – three genera, 33 species[164]
Genus name	Binomial authority	Classified	No. of species	Type species	Example image	Ref(s)
Acanthoponera	Mayr	1862	4	Acanthoponera mucronata	Acanthoponera mucronata	[64]
Aulacopone	Arnol'di	1930	1	Aulacopone relicta	Aulacopone relicta	[165]
Heteroponera	Mayr	1887	28	Heteroponera carinifrons	Heteroponera carinifrons	[80]

Leptanillinae

The subfamily Leptanillinae represents nine extant genera of ants, established in 1910 by Carlo Emery. They are subterranean ants from Africa, Europe and a single species known from Australia. Studies about their biology is minimal.^[166][167]

Subfamily Leptanillinae Emery, 1910 – nine genera, 64 species[168]
Genus name	Binomial authority	Classified	No. of species	Type species	Example image	Ref(s)
Anomalomyrma	Bolton	1990	3	Anomalomyrma taylori	Anomalomyrma taylori	[169]
Furcotanilla	Xu	2012	1	Furcotanilla furcomandibula	N/A	[170]
Leptanilla	Emery	1870	45	Leptanilla revelierii	Leptanilla revelierii	[171]
Noonilla	Petersen	1968	1	Noonilla copiosa	Noonilla copiosa	[172]
Opamyrma	Yamane, Bui & Eguchi	2008	1	Opamyrma hungvuong	Opamyrma hungvuong	[173]
Phaulomyrma	Wheeler & Wheeler	1930	1	Phaulomyrma javana	Phaulomyrma javana	[174]
Protanilla	Taylor	1990	9	Protanilla rafflesi	Protanilla rafflesi	[169]
Scyphodon	Brues	1925	1	Scyphodon anomalum	Scyphodon anomalum	[175]
Yavnella	Kugler	1987	2	Yavnella argamani	Yavnella argamani	[176]

Martialinae

The subfamily Martialinae contains the single genus Martialis which only has one species, Martialis heureka. The ant was discovered in 2000 Amazon rainforest near Manaus, Brazil. Described in 2008, the ant belongs to the oldest known distinct lineage to have diverged from the ancestors of all other ants.^[177]

Subfamily Martialinae Rabeling & Verhaagh, 2008 – one genus, one species[178]
Genus name	Binomial authority	Classified	No. of species	Type species	Example image	Ref(s)
Martialis	Rabeling & Verhaagh	2008	1	Martialis heureka	Martialis heureka	[177]

Myrmeciinae

The subfamily Myrmeciinae represents two extant genera and five fossil genera that were once found worldwide.^{[lower-alpha 4]} Established by Carlo Emery in 1877, the extant genera are restricted to Australia, New Caledonia and New Zealand.^[180] The notorious ant genus Myrmecia is known for their venomous stings and aggression, which has caused several human deaths in sensitive people.

Subfamily Myrmeciinae Emery, 1877 – seven genera, 111 species[181]
Genus name	Binomial authority	Classified	No. of species	Type species	Example image	Ref(s)
Archimyrmex !†Archimyrmex	Cockerell	1923	4	Archimyrmex rostratus !†Archimyrmex rostratus	†Archimyrmex rostratus	[182]
Avitomyrmex !†Avitomyrmex	Archibald, Cover & Moreau	2006	3	Avitomyrmex mastax !†Avitomyrmex mastax	N/A	[183]
Macabeemyrma !†Macabeemyrma	Archibald, Cover & Moreau	2006	1	Macabeemyrma ovata !†Macabeemyrma ovata	†Macabeemyrma ovata	[183]
Myrmecia	Fabricius	1804	95	Myrmecia gulosa	Myrmecia gulosa	[88][137]
Myrmeciites !†Myrmeciites[lower-alpha 4]	Archibald, Cover & Moreau	2006	0	None[lower-alpha 3]	†Myrmeciites "incertae sedis"	[183]
Nothomyrmecia	Clark	1934	1	Nothomyrmecia macrops	Nothomyrmecia macrops	[184]
Prionomyrmex !†Prionomyrmex	Mayr	1868	3	Prionomyrmex longiceps !†Prionomyrmex longiceps	†Prionomyrmex longiceps	[130]
Ypresiomyrma !†Ypresiomyrma	Archibald, Cover & Moreau	2006	4	Ypresiomyrma orbiculata !†Ypresiomyrma orbiculata	†Ypresiomyrma orbiculata	[183]

Myrmicinae

The subfamily Myrmicinae was established by Lepeletier de Saint-Fargeau in 1835. It represents 141 extant genera and 37 fossil genera that are distributed globally. It is the largest subfamily of the Formicidae, with more than 6,700 species described. The seed-harvesting ants and fungus-growing ants are well known among the Myrmicines.^[42][185]

Subfamily Myrmicinae Lepeletier de Saint-Fargeau, 1835 – 178 genera, 6,731 species[185]
Genus name	Binomial authority	Classified	No. of species	Type species	Example image	Ref(s)
Acanthognathus	Mayr	1887	8	Acanthognathus ocellatus	Acanthognathus ocellatus	[80]
Acanthomyrmex	Emery	1893	17	Acanthomyrmex luciolae	Acanthomyrmex concavus	[30]
Acromyrmex	Mayr	1865	32	Formica hystrix	Acromyrmex balzani	[186]
Adelomyrmex	Emery	1897	30	Adelomyrmex biroi	Adelomyrmex biroi	[187]
Adlerzia	Forel	1902	1	Monomorium froggatti	Adlerzia froggatti	[62]
Afromyrma !†Afromyrma	Dlussky, Brothers & Rasnitsyn	2004	1	Afromyrma petrosa !†Afromyrma petrosa	N/A	[39]
Agastomyrma !†Agastomyrma	Dlussky, Rasnitsyn & Perfilieva	2015	1	Agastomyrma laticeps !†Agastomyrma laticeps	N/A	[188]
Allomerus	Mayr	1878	8	Allomerus decemarticulatus	Allomerus decemarticulatus	[189]
Ancyridris	Wheeler	1935	2	Ancyridris polyrhachioides	Ancyridris polyrhachioides	[190]
Anillomyrma	Emery	1913	2	Monomorium decamerum	Anillomyrma decamera	[191]
Aphaenogaster	Mayr	1853	204	Aphaenogaster sardoa	Aphaenogaster swammerdami	[192]
Apterostigma	Mayr	1865	47	Apterostigma pilosum	Apterostigma pilosum	[186]
Aretidris	General	2015	2	Aretidris buenaventei	Aretidris buenaventei	[193]
Atopomyrmex	André	1889	3	Atopomyrmex mocquerysi	Atopomyrmex mocquerysi	[194]
Atta	Fabricius	1804	17	Formica cephalotes	Atta cephalotes	[137]
Attaichnus !†Attaichnus	Laza	1982	1	None	N/A	[195]
Attopsis !†Attopsis	Heer	1850	1	Attopsis longipennis !†Attopsis longipennis	N/A	[134]
Austromorium	Shattuck	2009	2	Xiphomyrmex flavigaster	Austromorium flavigaster	[196]
Baracidris	Bolton	1981	3	Baracidris meketra	Baracidris sitra	[197]
Bariamyrma	Lattke	1990	1	Bariamyrma hispidula	Bariamyrma hispidula	[198]
Basiceros	Schulz	1906	8	Meranoplus singularis	Basiceros manni	[199]
Biamomyrma !†Biamomyrma	Dlussky, Rasnitsyn & Perfilieva	2015	3	Biamomyrma zherikhini !†Biamomyrma zherikhini	N/A	[188]
Bilobomyrma !†Bilobomyrma	Radchenko & Dlussky	2013	2	Bilobomyrma ukrainica !†Bilobomyrma ukrainica	N/A	[200]
Blepharidatta	Wheeler	1915	4	Blepharidatta brasiliensis	Blepharidatta brasiliensis	[201]
Boltonidris !†Boltonidris	Radchenko & Dlussky	2012	1	Boltonidris mirabilis !†Boltonidris mirabilis	N/A	[202]
Bondroitia	Forel	1911	2	Monomorium coecum	Bondroitia lujae	[203]
Brachytarsites !†Brachytarsites	Hong	2002	1	Brachytarsites dongzhouheensis !†Brachytarsites dongzhouheensis	N/A	[125]
Calyptomyrmex	Emery	1887	37	Calyptomyrmex beccarii	Calyptomyrmex beccarii	[155]
Cardiocondyla	Emery	1869	70	Cardiocondyla elegans	Cardiocondyla wroughtonii	[204]
Carebara	Westwood	1840	212	Carebara lignata	Carebara longii	[205]
Cataulacus	Smith	1853	68	Cataulacus taprobanae	Cataulacus oberthueri	[206]
Cephalomyrmex !†Cephalomyrmex	Carpenter	1930	1	Cephalomyrmex rotundatus !†Cephalomyrmex rotundatus	†Cephalomyrmex rotundatus	[3]
Cephalotes	Latreille	1802	135	Formica atrata	Cephalotes atratus	[207]
Chimaeridris	Wilson	1989	2	Chimaeridris boltoni	Chimaeridris boltoni	[208]
Clavipetiola !†Clavipetiola	Hong	2002	1	Clavipetiola asiatica !†Clavipetiola asiatica	N/A	[125]
Colobostruma	Wheeler	1927	16	Epopostruma leae	Colobostruma foliacea	[209]
Crematogaster	Lund	1831	496	Formica scutellaris	Crematogaster scutellaris	[56]
Cryptomyrmex	Fernández	2004	2	Adelomyrmex longinodus	Cryptomyrmex longinodus	[210]
Cyatta	Sosa-Calvo et al.	2013	1	Cyatta abscondita	Cyatta abscondita	[211]
Cyphoidris	Weber	1952	4	Cyphoidris spinosa	Cyphoidris spinosa	[212]
Cyphomyrmex	Mayr	1862	41	Cyphomyrmex minutus	Cyphomyrmex minutus	[64]
Dacatria	Rigato	1994	1	Dacatria templaris	Dacatria templaris	[213]
Dacetinops	Brown & Wilson	1957	7	Dacetinops cibdelus	Dacetinops cibdelus	[214]
Daceton	Perty	1833	2	Formica armigera	Daceton armigerum	[215]
Diaphoromyrma	Fernández, Delabie & Nascimento	2009	1	Diaphoromyrma sofiae	Diaphoromyrma sofiae	[216]
Dicroaspis	Emery	1908	2	Dicroaspis cryptocera	Dicroaspis cryptocera	[217]
Dilobocondyla	Santschi	1910	19	Atopomyrmex selebensis	Dilobocondyla fouqueti	[218]
Diplomorium	Mayr	1901	1	Diplomorium longipenne	Diplomorium longipenne	[219]
Dolopomyrmex	Cover & Deyrup	2007	1	Dolopomyrmex pilatus	Dolopomyrmex pilatus	[220]
Electromyrmex !†Electromyrmex	Wheeler	1910	1	Electromyrmex klebsi !†Electromyrmex klebsi	†Electromyrmex klebsi	[221]
Enneamerus !†Enneamerus	Mayr	1868	1	Enneamerus reticulatus !†Enneamerus reticulatus	†Enneamerus reticulatus	[130]
Eocenidris !†Eocenidris	Wilson	1985	1	Eocenidris crassa !†Eocenidris crassa	†Eocenidris crassa	[154]
Eocenomyrma !†Eocenomyrma	Dlussky & Radchenko	2006	4	Eocenomyrma orthospina !†Eocenomyrma orthospina	†Eocenomyrma orthospina	[222]
Eomyrmex !†Eomyrmex	Hong	1974	1	Eomyrmex guchengziensis !†Eomyrmex guchengziensis	N/A	[223]
Epelysidris	Bolton	1987	1	Epelysidris brocha	Epelysidris brocha	[224]
Epopostruma	Forel	1895	19	Strumigenys quadrispinosa	Epopostruma natalae	[74]
Eurhopalothrix	Brown & Kempf	1961	53	Rhopalothrix bolaui	Eurhopalothrix bolaui	[225]
Eutetramorium	Emery	1899	3	Eutetramorium mocquerysi	Eutetramorium mocquerysi	[226]
Fallomyrma !†Fallomyrma	Dlussky & Radchenko	2006	1	Fallomyrma transversa !†Fallomyrma transversa	N/A	[227]
Formicoxenus	Mayr	1855	7	Myrmica nitidula	Formicoxenus nitidulus	[228]
Formosimyrma	Terayama	2009	1	Formosimyrma lanuyensis	Formosimyrma lanuyensis	[229]
Fushunomyrmex !†Fushunomyrmex	Hong	2002	1	Fushunomyrmex liaoningensis !†Fushunomyrmex liaoningensis	N/A	[125]
Gaoligongidris	Xu	2012	1	Gaoligongidris planodorsa	N/A	[230]
Gauromyrmex	Menozzi	1933	2	Gauromyrmex bengakalisi	Gauromyrmex bengakalisi	[231]
Goniomma	Emery	1895	8	Aphaenogaster blanci	Goniomma collingwoodi	[232]
Harpagoxenus	Forel	1893	3	Myrmica sublaevis	Harpagoxenus sublaevis	[233]
Huberia	Forel	1890	2	Tetramorium striatum	Huberia striata	[234]
Hylomyrma	Forel	1912	13	Pogonomyrmex columbicus	Hylomyrma balzani	[235]
Hypopomyrmex !†Hypopomyrmex	Emery	1891	1	Hypopomyrmex bombiccii !†Hypopomyrmex bombiccii	†Hypopomyrmex bombiccii	[236]
Ilemomyrmex !†Ilemomyrmex	Wilson	1985	1	Ilemomyrmex caecus !†Ilemomyrmex caecus	†Ilemomyrmex caecus	[237]
Indomyrma	Brown	1986	2	Indomyrma dasypyx	Indomyrma dasypyx	[238]
Ishakidris	Bolton	1984	1	Ishakidris ascitaspis	Ishakidris ascitaspis	[239]
Kalathomyrmex	Klingenberg & Brandão	2009	1	Myrmicocrypta emeryi	Kalathomyrmex emeryi	[240]
Kartidris	Bolton	1991	6	Kartidris nyos	Kartidris nyos	[241]
Kempfidris	Fernández, Feitosa & Lattke	2014	1	Monomorium inusuale	Kempfidris inusualis	[242]
Lachnomyrmex	Wheeler	1910	16	Lachnomyrmex scrobiculatus	Lachnomyrmex scrobiculatus	[6]
Lasiomyrma	Terayama & Yamane	2000	4	Lasiomyrma gedensis	Lasiomyrma gedensis	[243]
Lenomyrmex	Fernández & Palacio	1999	6	Lenomyrmex mandibularis	Lenomyrmex colwelli	[244]
Leptothorax	Mayr	1855	19	Formica acervorum	Leptothorax acervorum	[228]
Liomyrmex	Mayr	1865	1	Myrmica caeca	Liomyrmex gestroi	[186]
Lonchomyrmex !†Lonchomyrmex	Mayr	1867	2	Formica freyeri !†Lonchomyrmex freyeri	N/A	[245]
Lophomyrmex	Emery	1892	13	Oecodoma quadrispinosa	Lophomyrmex longicornis	[246]
Lordomyrma	Emery	1897	34	Lordomyrma furcifera	Lordomyrma furcifera	[187]
Malagidris	Bolton & Fisher	2014	6	Aphaenogaster belti	Malagidris belti	[247]
Manica	Jurine	1807	6	Formica rubida	Manica rubida	[89]
Mayriella	Forel	1902	9	Mayriella abstinens	Mayriella abstinens	[62]
Megalomyrmex	Forel	1885	44	Megalomyrmex leoninus	Megalomyrmex leoninus	[248]
Melissotarsus	Emery	1877	4	Melissotarsus beccarii	Melissotarsus beccarii	[249]
Meranoplus	Smith	1853	90	Cryptocerus bicolor	Meranoplus bicolor	[206]
Mesostruma	Brown	1948	9	Strumigenys turneri	Mesostruma turneri	[250]
Messor	Forel	1890	110	Formica barbara	Messor barbarus	[251]
Metapone	Forel	1911	18	Metapone greeni	Metapone emersoni	[252]
Microdaceton	Santschi	1913	4	Microdaceton exornatum	Microdaceton exornatum	[253]
Miosolenopsis !†Miosolenopsis	Zhang	1989	1	Miosolenopsis fossilis !†Miosolenopsis fossilis	N/A	[44]
Monomorium	Mayr	1855	372	Monomorium monomorium	Monomorium pharaonis	[228]
Mycetagroicus	Brandão & Mayhé-Nunes	2001	4	Mycetagroicus cerradensis	Mycetagroicus cerradensis	[254]
Mycetarotes	Emery	1913	4	Cyphomyrmex parallelus	Mycetarotes parallelus	[191]
Mycetophylax	Emery	1913	3	Myrmicocrypta brittoni	Mycetophylax conformis	[191]
Mycetosoritis	Wheeler	1907	5	Atta hartmanni	Mycetosoritis hartmanni	[255]
Mycocepurus	Forel	1893	6	Atta smithii	Mycocepurus smithii	[256]
Myrmecina	Curtis	1829	51	Myrmecina latreillii	Myrmecina graminicola	[257]
Myrmecites !†Myrmecites	Dlussky & Rasnitsyn	2003	6	Myrmecites rotundiceps !†Myrmecites rotundiceps	N/A	[258]
Myrmica	Latreille	1804	211	Formica rubra	Myrmica rubra	[87]
Myrmicaria	Saunders	1842	32	Myrmicaria brunnea	Myrmicaria brunnea	[259]
Myrmicocrypta	Smith	1860	27	Myrmicocrypta squamosa	Myrmicocrypta squamosa	[260]
Myrmisaraka	Bolton & Fisher	2014	2	Myrmisaraka producta	Myrmisaraka producta	[247]
Nesomyrmex	Wheeler	1910	57	Nesomyrmex clavipilis	Nesomyrmex angulatus	[6]
Novomessor	Emery	1915	3	Aphaenogaster cockerelli	Novomessor cockerelli	[261]
Ochetomyrmex	Mayr	1878	2	Ochetomyrmex semipolitus	Ochetomyrmex semipolitus	[189]
Octostruma	Forel	1912	34	Rhopalothrix simoni	Octostruma iheringi	[262]
Ocymyrmex	Emery	1886	37	Ocymyrmex barbiger	Ocymyrmex dekerus	[263]
Orbigastrula !†Orbigastrula	Hong	2002	1	Orbigastrula guchengziensis !†Orbigastrula guchengziensis	N/A	[125]
Orectognathus	Smith	1853	29	Orectognathus antennatus	Orectognathus antennatus	[206]
Oxyepoecus	Santschi	1926	21	Oxyepoecus bruchi	Oxyepoecus bruchi	[264]
Oxyidris !†Oxyidris	Wilson	1985	1	Oxyidris antillana !†Oxyidris antillana	†Oxyidris antillana	[237]
Oxyopomyrmex	André	1881	12	Oxyopomyrmex oculatus	Oxyopomyrmex oculatus	[265]
Parameranoplus !†Parameranoplus	Wheeler	1915	1	Parameranoplus primaevus !†Parameranoplus primaevus	†Parameranoplus primaevus	[7]
Paramycetophylax	Kusnezov	1956	1	Sericomyrmex bruchi	Paramycetophylax bruchi	[266]
Paraphaenogaster !†Paraphaenogaster	Dlussky	1981	4	Paraphaenogaster microphthalma !†Paraphaenogaster microphthalmus	†Paraphaenogaster hooleyana	[65]
Paratopula	Wheeler	1919	11	Atopomyrmex ceylonicus	Paratopula macta	[267]
Patagonomyrmex	Johnson & Moreau	2016	3	Pogonomyrmex angustus	Patagonomyrmex angustus	[268]
Perissomyrmex	Smith	1947	6	Perissomyrmex snyderi	Perissomyrmex snyderi	[269]
Peronomyrmex	Viehmeyer	1922	3	Peronomyrmex overbecki	Peronomyrmex greavesi	[270]
Phalacromyrmex	Kempf	1960	1	Phalacromyrmex fugax	Phalacromyrmex fugax	[271]
Pheidole	Westwood	1839	1007	Atta providens	Pheidole rhea	[272]
Pilotrochus	Brown	1978	1	Pilotrochus besmerus	Pilotrochus besmerus	[273]
Plesiomyrmex !†Plesiomyrmex	Dlussky & Radchenko	2009	1	Plesiomyrmex tubulatus !†Plesiomyrmex tubulatus	N/A	[274]
Podomyrma	Smith	1859	52	Podomyrma femorata	Podomyrma minor	[275]
Poecilomyrma	Mann	1921	2	Poecilomyrma senirewae	Poecilomyrma senirewae	[276]
Pogonomyrmex	Mayr	1868	69	Formica badia	Camponotus badius	[118]
Pristomyrmex	Mayr	1866	60	Pristomyrmex pungens	Pristomyrmex punctatus	[97]
Proatta	Forel	1912	1	Proatta butteli	Proatta butteli	[277]
Procryptocerus	Emery	1887	45	Meranoplus striatus	Procryptocerus hylaeus	[155]
Propodilobus	Branstetter	2009	1	Stenamma orientale	Propodilobus orientale	[278]
Protalaridris	Brown	1980	1	Protalaridris armata	Protalaridris armata	[279]
Protomyrmica !†Protomyrmica	Dlussky & Radchenko	2009	1	Protomyrmica atavia !†Protomyrmica atavia	†Protomyrmica atavia	[274]
Pseudoatta	Gallardo	1916	1	Pseudoatta argentina	Pseudoatta argentina	[280]
Quadrulicapito !†Quadrulicapito	Hong	2002	1	Quadrulicapito longa !†Quadrulicapito longa	N/A	[125]
Quineangulicapito !†Quineangulicapito	Hong	2002	1	Quineangulicapito fushunensis !†Quineangulicapito fushunensis	N/A	[125]
Recurvidris	Bolton	1992	11	None	Recurvidris recurvispinosa	[281]
Rhopalomastix	Forel	1900	6	Rhopalomastix rothneyi	Rhopalomastix rothneyi	[282]
Rhopalothrix	Mayr	1870	16	Rhopalothrix ciliata	Rhopalothrix ciliata	[283]
Rogeria	Emery	1894	40	Rogeria curvipubens	Rogeria curvipubens	[284]
Romblonella	Wheeler	1935	9	Romblonella grandinodis	Romblonella opaca	[190]
Rostromyrmex	Rosciszewski	1994	1	Rostromyrmex pasohensis	Rostromyrmex pasohensis	[285]
Rotastruma	Bolton	1991	2	Rotastruma recava	Rotastruma recava	[241]
Royidris	Bolton & Fisher	2014	15	Monomorium robertsoni	Royidris robertsoni	[247]
Secostruma	Bolton	1988	1	Secostruma lethifera	Secostruma lethifera	[286]
Sericomyrmex	Mayr	1865	19	Sericomyrmex opacus	Sericomyrmex amabilis	[186]
Sinomyrmex !†Sinomyrmex	Hong	2002	2	Sinomyrmex brunneus !†Sinomyrmex brunneus	N/A	[125]
Solenopsis	Westwood	1840	205	Solenopsis mandibularis	Solenopsis geminata	[205]
Solenopsites !†Solenopsites	Dlussky & Rasnitsyn	2003	3	Solenopsites minutus !†Solenopsites minutus	†Solenopsites rossi	[258]
Sphaerogasterites !†Sphaerogasterites	Hong	2002	1	Sphaerogasterites longipetiolatus !†Sphaerogasterites longipetiolatus	N/A	[125]
Stegomyrmex	Emery	1912	5	Stegomyrmex connectens	Stegomyrmex vizottoi	[287]
Stenamma	Westwood	1839	85	Stenamma westwoodii	Stenamma westwoodii	[272]
Stereomyrmex	Emery	1901	3	Stereomyrmex horni	Stereomyrmex horni	[288]
Stigmomyrmex !†Stigmomyrmex	Mayr	1868	1	Stigmomyrmex venustus !†Stigmomyrmex venustus	†Stigmomyrmex venustus	[130]
Stiphromyrmex !†Stiphromyrmex	Wheeler	1915	1	Stiphromyrmex robustus !†Stiphromyrmex robustus	†Stiphromyrmex robustus	[7]
Strongylognathus	Mayr	1853	24	Eciton testaceum	Strongylognathus testaceus	[289][290]
Strumigenys	Smith	1860	842	Strumigenys mandibularis	Strumigenys abdera	[260]
Syllophopsis	Santschi	1915	20	Monomorium modestum	Syllophopsis modesta	[291]
Talaridris	Weber	1941	1	Talaridris mandibularis	Talaridris mandibularis	[49]
Temnothorax	Mayr	1861	395	Myrmica recedens	Temnothorax unifasciatus	[68]
Terataner	Emery	1912	12	Atopomyrmex foreli	Terataner foreli	[287]
Tetheamyrma	Bolton	1991	1	Tetheamyrma subspongia	Tetheamyrma subspongia	[241]
Tetramorium	Mayr	1855	569	Formica caespitum	Tetramorium caespitum	[228]
Trachymyrmex	Forel	1893	48	Atta septentrionalis	Trachymyrmex septentrionalis	[256]
Tranopelta	Mayr	1866	2	Tranopelta gilva	Tranopelta gilva	[24]
Trichomyrmex	Mayr	1865	18	Trichomyrmex rogeri	Trichomyrmex destructor	[186]
Tropidomyrmex	Silva, Feitosa, Brandão & Diniz	2009	1	Tropidomyrmex elianae	Tropidomyrmex elianae	[292]
Tyrannomyrmex	Fernández	2003	3	Tyrannomyrmex rex	Tyrannomyrmex dux	[293]
Veromessor	Forel	1917	9	Aphaenogaster andrei	Veromessor andrei	[294]
Vitsika	Bolton & Fisher	2014	14	Vitsika crebra	Vitsika crebra	[247]
Vollenhovia	Mayr	1865	62	Vollenhovia punctatostriata	Vollenhovia emeryi	[186]
Vombisidris	Bolton	1991	17	Vombisidris philax	Vombisidris philax	[241]
Wasmannia	Forel	1893	11	Tetramorium auropunctatum	Wasmannia auropunctata	[295]
Wumyrmex !†Wumyrmex	Hong	2002	1	Wumyrmex furvis !†Wumyrmex furvis	N/A	[125]
Xenomyrmex	Forel	1885	4	Xenomyrmex stollii	Xenomyrmex stollii	[248]
Zhangidris !†Zhangidris	Bolton	2003	1	Zhangidris atopogaster !†Heteromyrmex atopogaster	N/A	[4]

Paraponerinae

The subfamily Paraponerinae contains a single genus Paraponera. This genus has two species, one of which was found in Dominican amber from the Miocene.^[296] The extant species, Paraponera clavata, is found in Central America and South America, and the pain from its sting is said to be greater than any other insect sting on earth.^[297]

Subfamily Paraponerinae Emery, 1901 – one genus, two species[298]
Genus name	Binomial authority	Classified	No. of species	Type species	Example image	Ref(s)
Paraponera	Smith	1858	2	Paraponera clavata	Paraponera clavata	[103]

Ponerinae

The subfamily Ponerinae was established by Lepeletier de Saint-Fargeau in 1835, which represents 47 extant genera and 12 fossil genera. The subfamily is among the most diverse in the family Formicidae, with more than 1,000 species described. They are mostly distributed in the tropics and subtropics.^[4][42]

Subfamily Ponerinae Lepeletier de Saint-Fargeau, 1835 – 59 genera, 1,287 species[299]
Genus name	Binomial authority	Classified	No. of species	Type species	Example image	Ref(s)
Afropone !†Afropone	Dlussky et al.	2004	2	Afropone oculata	N/A	[39]
Anochetus	Mayr	1861	122	Anochetus ghilianii	Anochetus ghilianii	[68]
Archiponera !†Archiponera	Carpenter	1930	1	Archiponera wheeleri !†Archiponera wheeleri	†Archiponera wheeleri	[3]
Asphinctopone	Santschi	1914	3	Asphinctopone silvestrii	Asphinctopone silvestrii	[26]
Austroponera	Schmidt & Shattuck	2014	3	Austroponera rufonigra	Austroponera rufonigra	[300]
Belonopelta	Mayr	1870	2	Belonopelta attenuata	Belonopelta attenuata	[283]
Boloponera	Fisher	2006	1	Boloponera vicans	Boloponera vicans	[301]
Bothroponera	Mayr	1862	37	Bothroponera pumicosa	Bothroponera pumicosa	[64]
Brachyponera	Emery	1900	19	Brachyponera croceicornis	Brachyponera croceicornis	[302]
Buniapone	Schmidt & Shattuck, 2014	2014	1	Buniapone amblyops	Buniapone amblyops	[300]
Centromyrmex	Mayr	1866	15	Centromyrmex bohemanni	Centromyrmex bohemanni	[97]
Cephalopone !†Cephalopone	Dlussky & Wedmann	2012	2	Cephalopone potens !†Cephalopone potens	†Cephalopone potens	[11]
Cryptopone	Emery	1893	24	Cryptopone testacea	Cryptopone testacea	[30]
Cyrtopone !†Cyrtopone	Dlussky & Wedmann	2012	4	†Cyrtopone microcephala	†Cyrtopone microcephala	[11]
Diacamma	Mayr	1862	24	Diacamma rugosum	Diacamma rugosum	[64]
Dinoponera	Roger	1861	8	Dinoponera gigantea	Dinoponera gigantea	[20]
Dolioponera	Brown	1974	1	Dolioponera fustigera	Dolioponera fustigera	[303]
Ectomomyrmex	Mayr	1867	28	Ectomomyrmex javanus	Ectomomyrmex javanus	[21][304]
Emeryopone	Forel	1912	5	Emeryopone buttelreepeni	Emeryopone buttelreepeni	[277]
Eogorgites !†Eogorgites	Hong	2002	1	Eogorgites cingulatus !†Eogorgites cingulatus	N/A	[125]
Eoponerites !†Eoponerites	Hong	2002	1	Eoponerites longipetiolatus !†Eoponerites longipetiolatus	N/A	[125]
Euponera	Forel	1891	27	Euponera sikorae	Euponera sikorae	[96]
Feroponera	Bolton & Fisher	2008	1	Feroponera ferox	Feroponera ferox	[305]
Fisheropone	Schmidt & Shattuck	2014	1	Fisheropone ambigua	Fisheropone ambigua	[300]
Furcisutura !†Furcisutura	Hong	2002	1	Furcisutura wanghuacunensis !†Furcisutura wanghuacunensis	N/A	[125]
Hagensia	Forel	1901	2	Hagensia havilandi	Hagensia havilandi	[306]
Harpegnathos	Jerdon	1851	7	Harpegnathos saltator	Harpegnathos saltator	[307]
Hypoponera	Santschi	1938	154	Hypoponera abeillei	Hypoponera abeillei	[308]
Iroponera	Schmidt & Shattuck	2014	1	Iroponera odax	Iroponera odax	[300]
Leptogenys	Roger	1861	308	Leptogenys falcigera	Leptogenys falcigera	[20]
Loboponera	Bolton & Brown	2002	9	Loboponera vigilans	Loboponera vigilans	[309]
Longicapitia !†Longicapitia	Hong	2002	1	Longicapitia reticulata !†Longicapitia reticulata	N/A	[125]
Mayaponera	Schmidt & Shattuck	2014	1	Mayaponera constricta	Mayaponera constricta	[300]
Megaponera	Mayr	1862	1	Megaponera analis	Megaponera analis	[64]
Mesoponera	Emery	1900	20	Mesoponera melanaria	Mesoponera melanaria	[310]
Messelepone !†Messelepone	Dlussky & Wedmann	2012	1	†Messelepone leptogenoides	†Messelepone leptogenoides	[11]
Myopias	Roger	1861	40	Myopias amblyops	Myopias amblyops	[20]
Neoponera	Emery	1901	57	Neoponera villosa	Neoponera villosa	[311]
Odontomachus	Latreille	1804	70	Odontomachus haematodus	Odontomachus haematodus	[87]
Odontoponera	Mayr	1862	2	Odontoponera denticulata	Odontoponera denticulata	[64]
Ophthalmopone	Forel	1890	5	Ophthalmopone berthoudi	Ophthalmopone berthoudi	[234]
Pachycondyla	Smith	1858	36	Pachycondyla crassinoda	Pachycondyla crassinoda	[103]
Paltothyreus	Mayr	1862	1	Paltothyreus tarsatus	Paltothyreus tarsatus	[311]
Parvaponera	Schmidt & Shattuck	2014	4	Parvaponera darwinii	Parvaponera darwinii	[300]
Phrynoponera	Wheeler	1920	5	Phrynoponera gabonensis	Phrynoponera gabonensis	[124]
Platythyrea	Roger	1863	44	Platythyrea punctata	Platythyrea punctata	[21][104]
Plectroctena	Smith	1858	17	Plectroctena mandibularis	Plectroctena mandibularis	[103]
Ponera	Latreille	1804	62	Ponera coarctata	Ponera coarctata	[87][272]
Ponerites !†Ponerites	Dlussky & Rasnitsyn	2003	14	Ponerites eocenicus !†Ponerites eocenicus	†Ponerites eocenicus	[258]
Promyopias	Santschi	1914	1	Promyopias silvestrii	Promyopias silvestrii	[26]
Protopone !†Protopone	Dlussky	1988	7	Protopone primigena !†Protopone primigena	†Protopone primigena	[29]
Psalidomyrmex	André	1890	6	Psalidomyrmex foveolatus	Psalidomyrmex foveolatus	[312]
Pseudoneoponera	Donisthorpe	1943	18	Pseudoneoponera verecundae	Pseudoneoponera verecundae	[313]
Pseudoponera	Emery	1900	6	Pseudoponera stigma	Pseudoponera stigma	[302]
Rasopone	Schmidt & Shattuck	2014	11	Rasopone ferruginea	Rasopone ferruginea	[300]
Simopelta	Mann	1922	22	Simopelta jeckylli	Simopelta jeckylli	[314]
Streblognathus	Mayr	1862	2	Streblognathus aethiopicus	Streblognathus aethiopicus	[64]
Taphopone !†Taphopone	Dlussky & Perfilieva	2014	6	Taphopone karaganensis !†Taphopone karaganensis	†Taphopone macroptera	[31]
Thaumatomyrmex	Mayr	1887	12	Thaumatomyrmex mutilatus	Thaumatomyrmex mutilatus	[80]

Proceratiinae

The subfamily Proceratiinae was established by Italian entomologist Carlo Emery in 1895, which represents three extant genera and one extinct genus. Found worldwide, these ants are mainly encountered in tropical and subtropical areas. Little is known about their biology.^[42][315]

Subfamily Proceratiinae Emery, 1895 – four genera, 155 species[316]
Genus name	Binomial authority	Classified	No. of species	Type species	Example image	Ref(s)
Bradoponera !†Bradoponera	Mayr	1868	4	Bradoponera meieri !†Bradoponera meieri	†Bradoponera meieri	[130]
Discothyrea	Roger	1863	37	Discothyrea testacea	Discothyrea testacea	[104]
Probolomyrmex	Mayr	1901	26	Probolomyrmex filiformis	Probolomyrmex filiformis	[219]
Proceratium	Roger	1863	88	Proceratium silaceum	Proceratium silaceum	[104]

Pseudomyrmecinae

The subfamily Pseudomyrmecinae was established by M.R. Smith in 1952,^[317] which represents three genera of ants that are primarily arboreal nesting ants in the tropical and subtropical regions.^[318] They are found in Africa, Asia, Australia, North America and South America.^[319]

Subfamily Pseudomyrmecinae Smith, 1952 – three genera, 251 species[320]
Genus name	Binomial authority	Classified	No. of species	Type species	Example image	Ref(s)
Myrcidris	Ward	1990	1	Myrcidris epicharis	Myrcidris epicharis	[318]
Pseudomyrmex	Lund	1831	147	Pseudomyrmex gracilis	Pseudomyrmex gracilis	[317][321]
Tetraponera	Smith	1852	103	Tetraponera nigra	Tetraponera nigra	[322]

Sphecomyrminae

The subfamily Sphecomyrminae contains nine fossil genera of stem ants. Most fossilised ants from Cretaceous amber are placed in this subfamily.^[35][323]

Subfamily Sphecomyrminae Wilson & Brown, 1967 – nine genera, 18 species[324]
Genus name	Binomial authority	Classified	No. of species	Type species	Example image	Ref(s)
Baikuris !†Baikuris	Dlussky	1987	4	Baikuris mandibularis !†Baikuris mandibularis	†Baikuris mandibularis	[325]
Ceratomyrmex !†Ceratomyrmex	Perrichot, Wang & Engel	2016	1	Ceratomyrmex ellenbergeri !†Ceratomyrmex ellenbergeri	†Ceratomyrmex ellenbergeri	[326]
Cretomyrma !†Cretomyrma	Dlussky	1975	2	Cretomyrma arnoldii !†Cretomyrma arnoldii	N/A	[36]
Dlusskyidris !†Dlusskyidris	Bolton	1994	1	Dlusskyidris zherichini !†Dlusskyidris zherichini	†Dlusskyidris zherichini	[36][327]
†Haidomyrmex	Dlussky	1996	3	Haidomyrmex cerberus !†Haidomyrmex cerberus	†Haidomyrmex cerberus	[32]
Haidomyrmodes !†Haidomyrmodes	Perrichot et al.	2008	1	Haidomyrmodes mammuthus !†Haidomyrmodes mammuthus	†Haidomyrmodes mammuthus	[328]
Haidoterminus !†Haidoterminus	McKellar, Glasier & Engel	2013	1	Haidoterminus cippus !†Haidoterminus cippus	†Haidoterminus cippus	[329]
Sphecomyrma !†Sphecomyrma	Wilson & Brown	1967	3	Sphecomyrma freyi !†Sphecomyrma freyi	†Sphecomyrma freyi	[330]
Zigrasimecia !†Zigrasimecia	Barden & Grimaldi	2013	2	Zigrasimecia tonsora !†Zigrasimecia tonsora	†Zigrasimecia tonsora	[323]

Incertae sedis

There are several ant genera where their taxonomic placement is uncertain (incertae sedis). These genera have not yet been assigned to any subfamily within Formicidae; 14 genera are currently listed as incertae sedis.^[331]

Incertae sedis taxa
Genus name	Binomial authority	Classified	No. of species	Type species	Example image	Ref(s)
Calyptites !†Calyptites	Scudder	1877	1	Calyptites antediluvianum !†Calyptites antediluvianum	N/A	[3][332]
Camelomecia !†Camelomecia	Barden & Grimaldi	2016	1	Camelomecia janovitzi !†Camelomecia janovitzi	N/A	[333]
Condylodon	Lund	1831	1	Condylodon audouini	N/A	[4][56]
Cretopone !†Cretopone	Dlussky	1975	1	Cretopone magna !†Cretopone magna	†Cretopone magna	[334]
Curticorna !†Curticorna[lower-alpha 5]	Hong	2002	1	Curticorna leptogastrosa !†Curticorna leptogastrosa	N/A	[4][125]
Eoaenictites !†Eoaenictites[lower-alpha 6]	Hong	2002	1	Eoaenictites castanifurvus !†Eoaenictites castanifurvus	N/A	[125]
Eoformica !†Eoformica	Cockerell	1921	6	Eoformica pinguis !†Eoformica eocenica	†Eoformica pinguis	[258][337]
Fonsecahymen !†Fonsecahymen	Martins-Neto & Mendes	2002	1	Fonsecahymen stigmata !†Fonsecahymen stigmata	N/A	[338]
Gerontoformica !†Gerontoformica	Nel & Perrault	2004	13	Gerontoformica cretacica !†Gerontoformica cretacica	†Gerontoformica cretacica	[339]
Hypochira[lower-alpha 7]	Buckley	1866	1	Formica subspinosa	N/A	[340][341]
Klondikia !†Klondikia	Dlussky & Rasnitsyn	2003	1	Klondikia whiteae !†Klondikia whiteae	N/A	[258]
Kohlsimyrma !†Kohlsimyrma	Dlussky & Rasnitsyn	2003	3	Kohlsimyrma laticeps !†Kohlsimyrma laticeps	N/A	[258]
Myanmyrma !†Myanmyrma	Engel & Grimaldi	2005	1	Myanmyrma gracilis !†Myanmyrma gracilis	†Myanmyrma gracilis	[33]
Petropone !†Petropone	Dlussky	1975	1	Petropone petiolata !†Petropone petiolata	†Petropone petiolata	[334]

Excluded from Formicidae

Listed taxa were once believed to be ants and were placed within Formicidae, but at present they are no longer considered ants.

Taxa excluded from Formicidae
Genus name	Binomial authority	Classified	No. of species	Type species	Notes	Ref(s)
Cariridris !†Cariridris	Brandão & Martins-Neto	1990	1	†Cariridris bipetiolata	The fossil was thought to be a member of Myrmeciinae from the Cretaceous, but it is now incertae sedis in the subclade Aculeata.	[34][342]
Cretacoformica !†Cretacoformica	Jell & Duncan	1986	1	†Cretacoformica explicata	The fossil is an insect from the Lower Cretaceous in Australia that was incertae sedis in Formicidae, but it is now incertae sedis in the suborder Apocrita.	[34][343]
Formila !†Formila	Romand	1846	1	†Formila chevrolatii	Transferred to the wasp family Embolemidae, now incertae sedis in the suborder Symphyta.	[4][344]
†Myrmicium	Westwood	1854	1	†Myrmicium heerii	Transferred to Myrmicinae in 1893, but it is now placed in the sawfly family Embolemidae.	[110][145]
Promyrmicium !†Promyrmicium[lower-alpha 8]	Baroni Urbani	1971	1	None	Originally placed in Myrmicinae, now assigned to the family Ichneumonidae.	[4][345]
Syntaphus !†Syntaphus	Donisthorpe	1920	1	†Syntaphus wheeleri	The fossil is an insect from the Oligocene, originally placed in Ponerinae and later assigned to incertae sedis in Formicidae. It was recently excluded from the family in 2014, now believed to be a Braconidae wasp.	[4][31][59]

See also

• List of ant subfamilies

Notes

References